The gay linklists you mention are like a blast from the past - 1999 rules - it's been wondered many times why they still have the rules they do.
I've subbed sites to you and the other 'big names' in the gay linklist world - and oddly enough, gotten better ratios subbing smaller sites to the straight linklists. ( But maybe I built too clean. )
That could be just a coincidence - but I don't like building sites with 48-50-60 pics to get ratios that are the same or less as sites with 24 pics.
Here's a 'straight linklist' gay site that just made a sale today. Not my best niche, so I'm willing to share it. Not the greatest site...
http://www.gay-interracial-xxx.com/redhead-jarhead/
Yes, those sites you showed would be considered too clean for good sales by the standards of the 50pic gay linklists. Kind of similar to sites I subbed there.