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I'd guess that as long as the images are on the same domain, you could link to them. Dont take my word for it though. The linklist owners make the decisions.
I know they dont have to be in a single directory because I've had galleries listed that used seperate image, recip and css directories, but all three were subdirectories of the directory I submitted. It just helped keep things cleaner. What I've found though, is that in the end, its really just easier to put everything in one directory. Disk space is cheap.
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Its just a jump to the left.
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